Jakob Johann Freiherr von Uexküll (; German: [ˈʏkskʏl]; 8 September [O.S. 27 August] 1864 – 25 July 1944) was a Baltic German biologist who worked in the fields of muscular physiology and animal behaviour studies and was an influence on the cybernetics of life. However, his most notable contribution is the notion of Umwelt, used by semiotician Thomas Sebeok and philosopher Martin Heidegger. His works established biosemiotics as a field of research.
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