Leland Harrison "Lee" Hartwell (born October 30, 1939) is an American former president and director of the Fred Hutchinson Cancer Research Center in Seattle, Washington. He shared the 2001 Nobel Prize in Physiology or Medicine with Paul Nurse and Tim Hunt, for their discoveries of protein molecules that control the division (duplication) of cells.
Working in yeast, Hartwell identified the fundamental role of checkpoints in cell cycle control, and CDC genes such as CDC28, which controls the start of the cycle—the progression through G1.
HistoryLayer draws the world as it was in any year from 123,000 BCE to
2026. Move the slider and the borders redraw for that year; the map
carries 23,890 events, 12,790 people, 837 wars and 13,183 cities, each
one readable in place. Signing in is optional and lets you keep what
you find.